By Helen Bushby Entertainment and arts reporter

Image copyright Getty Images

Lady Gaga led this year's MTV VMAs, winning five awards, while BTS and Ariana Grande won four each and The Weeknd won video of the year.

The ceremony, filmed at several outdoor New York locations with a limited audience, was dedicated to the late Black Panther star Chadwick Boseman.

It included pre-recorded speeches and performances, with Lady Gaga saying: "Love who you are. Be kind. Mask up."

Host Keke Palmer called Boseman, who died on Friday, "a true hero".

Here are some key moments from the socially-distanced show, which was broadcast on Sunday night, with the full list of winners here.

1. Lady Gaga on 'wrath' and 'rage' of art

Image copyright Getty Images

Gaga's wins were for best artist, song, collaboration and cinematography, plus the first MTV Tricon Award - honouring her work in music, acting and fashion.

"This has not been an easy year for a lot of people, but what I see in the world is a massive triumph of courage," she said.

"Just because we're separated right now and culture may feel less alive in some ways, I know a renaissance is coming, and the wrath of pop culture will inspire you and the rage of art will empower you as it responds to hardship with its generosity and love. "

Model Bella Hadid, who called Gaga a "powerhouse" as she introduced her, said: "She turns pop music into high art. She redefines fashion.. she inspires and empowers all of us."

2. MTV remembers Chadwick Boseman

Host Keke Palmer said of Chadwick Boseman: "We dedicate tonight's show to a man whose spirit touched so many. He's a true hero, not just on screen but in everything he did. His impact lives forever."

The awards honoured the late actor, who died of colon cancer aged just 43, with a throwback to his 2018 appearance at the MTV Movie Awards, when he won best hero and best performance in a film for playing T'Challa in Black Panther.

During his speech, he invited James Shaw Jr to the stage to thank him for protecting other restaurant-goers from a gunman who opened fire in a waffle house in Tennessee.

He gave Shaw his popcorn trophy and said: "This is gonna live at your house. God bless you, man."

The clip of Boseman ended with the words "rest in power".

3. The Weeknd called for justice over shootings

The Weeknd won the highly prized video of the year award for Blinding Lights, using his speech to demand justice in the cases of two African Americans shot by police.

"It's hard to celebrate, so I'm just going to say justice for Jacob Blake and justice for Breonna Taylor," he said.

Da Baby's performance echoed this, referencing the ongoing protests against police violence, and singing his song Blind while handcuffed from the back of a police car.