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Los Angeles is offering free coronavirus tests to all residents, regardless of whether they are displaying any symptoms.
Mayor Eric Garcetti made the announcement just hours after Los Angeles County reported its largest daily increase in new Covid-19 cases.
Until now, only essential workers and those displaying symptoms could receive tests due to a scarcity of kits.
It comes as California is expected to close all beaches across the state.
Los Angeles County currently accounts for almost half of California's confirmed coronavirus cases. The state has so far reported over 48,000 cases and more than 1,900 deaths.
On Thursday, Mr Garcetti urged all of the city's residents to get swabbed after earlier tweeting that "LA is now the first major city in America to offer free Covid-19 testing".
Meanwhile, California Governor Gavin Newsom is expected to sign an executive order to close all beaches and parks in the state after large crowds packed public spaces over the weekend.
While some of California's beaches have been closed for weeks, others have remained open with social distancing rules in place.
Mr Newsom recently said that he was just weeks away from lifting some restrictions.
But the recent images of crowded beaches in the most populous US state led to a warning from the Democratic governor that the behaviour of some Californians threatened to undo weeks of work to restrict the spread of Covid-19.
"We can't see images like we saw, particularly on Saturday, in Newport Beach," the governor said on Monday, adding: "The virus doesn't take the weekend off because it's a beautiful sunny day around our coasts."
He later said, however, that he would not be making changes to orders "on the basis of pressure".
"What we need to do, from my humble perspective, is listen to the public health experts."
California was among the first US states to bring in blanket restrictions, issuing a "stay at home" order to residents last month in an effort to stem the march of the coronavirus.
By Sophie Long, BBC News, Los Angeles
Los Angeles Mayor Eric Garcetti's announcement of free testing for all came hours after a grim milestone - LA County saw it's largest increase in new Coronavirus cases reported in a single day since the pandemic began, though the rate of hospitalisations and percentage of people testing positive have remained stable.
Eric Garcetti urged all Angelenos to get swabbed. "You can't put a price on the peace of mind of knowing you can't infect someone around you" he said. California's leaders have long characterised expanding Covid-19 testing as one of a number of critical milestones that need to be met before stay at home orders can be modified.
While Los Angeles County, which is home to nearly 10 million people, continues to be the coronavirus hotbed of California, accounting for nearly half of the States' nearly 50,000 confirmed cases, that milestone could now be in sight.
In other US developments:
- US-wide social distancing measures expire on Thursday, US President Donald Trump has said he will not renew them
- More than one in six nursing homes in the US have recorded coronavirus cases, according to the Washington Post
- Georgia is to let 16-year-olds drive without a road test due to social distancing measures
- In the past week, 3.8 million more people filed for unemployment benefits in the US, meaning at least 30 million Americans have lost their jobs since mid-March
- Mr Trump stepped up his criticism of China over the outbreak, saying the country "will do anything they can" to make him lose in his re-election bid