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Venezuela's President Nicolás Maduro has announced he will be asking for volunteers to test the Covid-19 vaccine developed by Russia.

Russia was the first country to officially register a vaccine against the coronavirus on 11 August.

The Venezuelan government, which has received billions of dollars in loans from Russia, said it would be willing to take part in clinical trials.

"In the coming days, we'll be asking for volunteers to get vaccinated," President Maduro announced on Sunday.

Russian scientists had said early-stage trials of their vaccine were a success - although the vaccine's approval by regulators comes before the completion of a larger study involving thousands of people.

Mr Maduro's statement indicates that Venezuela does not have the volunteers yet, suggesting that Health Minister Carlos Alvarado jumped the gun when he said 10 days ago that Venezuela had offered Russia "500 volunteers to participate in vaccine trials".

The Venezuelan government has also said that it was ready to participate in testing of Chinese and Cuban vaccines against Covid-19.

China and Cuba, along with Russia, are staunch allies of Nicolás Maduro, whose government has been increasingly isolated after more than 50 nations, including the US and UK, have recognised his rival Juan Guaidó as the interim president.

Mr Guaidó, who leads the National Assembly - the only institution in Venezuela to be controlled by the opposition - declared himself acting president in January 2019 after the election which had returned Mr Maduro to power was widely deemed to be neither free nor fair.

Mr Maduro, who has the support of the security forces, has remained in the presidential palace and largely in control of the country, but US sanctions have left his government reliant on loans from Russia and his other remaining allies.

After Russia announced it had registered the Covid-19 vaccine, which it has named Sputnik V, Mr Maduro congratulated it on its achievement.

He also said on that he would be the first to get inoculated. "The first one to be vaccinated will be me. I'm going to get the vaccine, I'm going to set an example," he said on 16 August.

However, speaking on Sunday, he did not say whether he would be among the 500 Venezuelan volunteers.

Venezuela has fewer than 50,000 registered cases of coronavirus and 381 deaths, a much lower toll than other countries in the region.

But immunologists say the true figure could be much higher as many Venezuelans do not get tested for fear of being stigmatised after the government blamed returning migrants for the spread of Covid-19.